摘要
造孢细胞具有丰富的细胞器,胞间连丝明显。花粉母细胞中除有大量的细胞器外,还观察到由粗糙内质网环绕成的结构——同心圆内质网。它们形态各异,单独或成群分布,有的中央还环绕着淀粉粒。这在松属植物花粉母细胞中还未见有报导。此时期的绒毡层细胞内具大量成束聚集平行排列的内质网。四分体时期,绒毡层细胞通过质膜和内平周壁向药囊内分泌前乌氏体和孢粉素颗粒。生殖细胞与管细胞由一月牙形薄壁分隔,此两细胞在位置,形态以及内含物上均存在着差异。这决定了它们在雄配子体进一步发育中;有着不同的发育途径和功能。 本世纪六十年以后,对松属植物小孢子发生及花粉形态结构等方面的电子显微镜研究已积累了一些资料。有关云南松胚胎学方面的超微结构研究除刘成运等(1986)对云南松小孢子发生的超微结构进行了研究外,对其小孢子整个发生发育过程的超微结构变化的研究,至今未见有详细报道。因此,对这方面的系统研究是具有一定的理论意义。
This paper reports the observations on the ulstrastructure of microsp-orogenesis and development of Pinus yunnanesis Franch. There were a lot of cell-organelles in sporogenous cells and pollenmother cells. Plasmodesma were obvious within the sporogenous cell walls.Concentric circle-shaped endoplasmic reticula in the pollen mother cellswere observed. Starch was surrounded by this kind endoplasmic reticula.The concentric circle-shaped endoplasmic reticula appear variously anddistributed singly or concentrically in the pollen mother cells. During thisstage, many sharply-defined endoplasmic reticulum demonstrated in a para-llelly arranged bundle-shaped structure in the tapetum cells. At the tetradstage, pre-Ubisch bodies and sporopollenin grains from tapetum cells secr-eted into microsporangium through plasmalemma and inner tangential wall.Generative cell and tube cell were seperated by a mooned thin wall. Therewere differences in the position, shape, size and contents of the generativecell and tube cell. These differences decided the different development ways and functions of the generative cell and tube cell in the further dev-elopment of male gametophyte.
出处
《植物研究》
CAS
CSCD
北大核心
1989年第1期87-93,共7页
Bulletin of Botanical Research
