基金Acknowledgements This work was supported by the National Natural Sciences Foundation of China （NSFC 31272268/ 31172113）, the Special Foundation of Ministry of Science and Technology of the People＇ s Republic of China （MOST grant no. 2014FY110100） and the Natural Science Foundation of Guizhou Province （J [2014] 2146）. We are grateful to, Jianyong Li, Ziqi Li, Yaqian Fu and, Mu Yan, for collecting specimens. This work was supported by the National Natural Sciences Foundation of China （NSFC-31272268/31172113）, the Special Foundation of Ministry of Science and Technology of the People＇s Republic of China （MOST grant no. 2014FY 110100） and the Natural Science Foundation of Guizhou Province （J [2014] 2146）.
基金Acknowledgements We are deeply grateful to Leng Cong and Long Ping for collecting specimens. This work was supported by the National Natural Sciences Foundation of China （Grant No. 31172113 / 31272268 / 31601882）, the Special Foundation of Ministry of Science and Technology of the People＇ s Republic of China （Grant No. 2014FY 110100） and the Natural Science Foundation of Guizhou Province （Grant No. J [2014] 2146）.
基金This work was supported by the grants fi：om the National Natural Sciences Foundation of China （NSFC-31572276 and NS FC-31601850）, the Hunan Provincial Natural Science Foundation （2017JJ3202） and the Singapore Ministry of Education AcRF Tier 1 grant （R-154-000-A52-114）
