基金the Chinese Academy of Sciences Visiting Professorship Scheme(2011T1S20)the National Natural Science Foundation of China(31000258,31470306,31260038,31460159)+3 种基金the Key Laboratory of Plant Diversity and Biogeography of East AsiaKunming Institute of Botany(KIB)&the Chinese Academy of Sciences(2014CB954100)Guangxi Natural Science Foundation(2015GXNSFBB139004&2015GXNSFBA139105)Guangxi Forest Science&Technology Foundation(Gui Lin Ke Zi〔2014〕No.27)