基金Acknowledgements We are deeply grateful to Leng Cong and Long Ping for collecting specimens. This work was supported by the National Natural Sciences Foundation of China （Grant No. 31172113 / 31272268 / 31601882）, the Special Foundation of Ministry of Science and Technology of the People＇ s Republic of China （Grant No. 2014FY 110100） and the Natural Science Foundation of Guizhou Province （Grant No. J [2014] 2146）.
