基金supported by the National Natural Science Foundation of China(U22A20561 and 41922005)the Tibetan Plateau Scientific Expedition and Research Program(2019QZKK0202)+3 种基金the National Key Research and Development Program of China(2022YFC3204100)the Nanjing Institute of Geography and Limnology,Chinese Academy of Sciences Foundation(E1SL002)supported by the Tübitak Program BIDEB2232(Project 118C250)supported by a UK Research and Innovation(UKRI)Natural Environment Research Council(NERC)Independent Research Fellowship(NE/T011246/1)。