基金supported by the National Natural Science Foundation of China(42030508,41988101)the Second Tibetan Plateau Scientific Expedition and Research Program(STEP)(2019QZKK0301)+1 种基金funding from the European Research Council(ERC-SyG-2013-610028 IMBALANCE-P)funding from the project “Inside out”(#POIR.04.04.00-00-5F85/18-00)funded by the HOMING programme of the Foundation for Polish Science co-financed by the European Union under the European Regional Development Fund。