基金国家自然科学基金项目(Nos.40930847,31070442)福建省自然科学基金项目(Nos.2012J01150,2010J01223)+2 种基金福建省教育厅项目(No.JA10232)莆田市科技计划项目[No.2011S09(4)]资助the Open Fund of the Key Laboratory of the Ministry of Education for Coastal and Wetland Ecosystems of Xiamen University(CWel0902)~~