基金supported from the National Key Research and Development Program(2021YFF0502300)the National Natural Science Foundation of China(42325603,42022046,42227803,and 41890850)+1 种基金the Guangdong Natural Resources Foundation(GDNRC[2022]45)the PI Project of Southern Marine Science and Engineering Guangdong Laboratory(Guangzhou)(GML20190609 and GML2022009)。