基金supported by the Strategic Priority Research Program of the Chinese Academy of Sciences(XDA13010101)the Key Research and Development Plan of Hainan Province(ZDYF2020198)+3 种基金the Guangdong Research Foundation(2019BT02H594)the National Natural Science Foundation of China(42076071)the Key Special Project for Introduced Talents Team of the Southern Marine Science and Engineering Guangdong Laboratory(Guangzhou)(GML2019ZD0204)the Rising Star Foundation of the South China Sea Institute Oceanology(NHXX2017DZ0101)。