The Triassic rocks are widespread in China, and both marine and terrestrial strata are well developed. The Triassic stratigraphic architecture of China is very complex in both spatial variation of the so-called "...The Triassic rocks are widespread in China, and both marine and terrestrial strata are well developed. The Triassic stratigraphic architecture of China is very complex in both spatial variation of the so-called "South Marine and North Continental", i.e. the southern areas of China occupied mostly by marine facies while the northern China by terrestrial facies during the Triassic Period, and temporal transition of the "Lower Marine and Upper Continental", i.e. the lower part of the Triassic System composed mainly of marine facies and the upper part of terrestrial strata especially in South China. Although the Global Stratotype Section and Point(GSSP) of the Permian-Triassic boundary is located in South China, the Triassic of China except for some marine Lower-Middle Triassic depositions shows significantly local characteristics and is hardly correlated with the global chronostratigraphic chart. Consequently, the Triassic of China contains not only the international research hotspots but also difficult points in stratigraphic study. This paper aims to present a brief review of the Triassic in China, including chronostratigraphy, biostratigraphy, magnetostratigraphy and chemostratigraphy, and summarize an integrated Triassic stratigraphic framework of China. Accordingly, a stratigraphic correlation is proposed for the lithostratigraphic sequences among the three tectono-paleogeographic stratigraphic regions. The comprehensive study indicates that ammonoids are the classic index fossils in Triassic biostratigraphy but conodonts are more advantageous in the study and definition of the Triassic chronostratigraphic boundaries. China still has the potential to optimize the GSSPs of the Induan-Olenekian boundary and Olenekian-Anisian boundary. The correlation of the Permian-Triassic boundary between marine and terrestrial facies might be achieved with the help of the Permian-Triassic "transitional bed" and its related biotic and environmental events in association with the biostratigraphic study of conchostracan, vertebra展开更多
Ocean temperature and dissolved oxygen concentrations are critical factors that control ocean productivity, carbon and nutrient cycles, and marine habitat. However, the evolution of these two factors in the geologic p...Ocean temperature and dissolved oxygen concentrations are critical factors that control ocean productivity, carbon and nutrient cycles, and marine habitat. However, the evolution of these two factors in the geologic past are still unclear. Here, we use a new oxygen isotope database to establish the sea surface temperature(SST) curve in the past 500 million years. The database is composed of 22 796 oxygen isotope values of phosphatic and calcareous fossils. The result shows two prolonged cooling events happened in the Late Paleozoic and Late Cenozoic, coinciding with two major ice ages indicated by continental glaciation data, and seven global warming events that happened in the Late Cambrian, Silurian–Devonian transition, Late Devonian, Early Triassic, Toarcian, Late Cretaceous, and Paleocene–Eocene transition. The SSTs during these warming periods are about 5–30 °C higher than the present-day level. Oxygen contents of shallow seawater are calculated from temperature, salinity, and atmospheric oxygen. The results show that major dissolved oxygen valleys of surface seawater coincide with global warming events and ocean anoxic events. We propose that the combined effect of temperature and dissolved oxygen account for the long-term evolution of global oceanic redox state during the Phanerozoic.展开更多
The miniaturization of organisms during the Permian-Triassic mass extinction,as an ecological strategy in response to environmental devastation,has been widely recognized in diverse marine invertebrates.Previous studi...The miniaturization of organisms during the Permian-Triassic mass extinction,as an ecological strategy in response to environmental devastation,has been widely recognized in diverse marine invertebrates.Previous studies on the extinction process and miniaturization of foraminifers in the Permian-Triassic interval have relied on the fossil record of the low-latitude Paleotethys or a global database,although data and materials from the high-latitude Neotethys region are still rare.To reveal the evolutionary patterns and spatial variability of foraminifers at different latitudes and paleogeographic contexts,here we investigated the fossil distribution and size variation of foraminifers in the Selong Section of southern Tibet,located in the mid-latitude Neotethys of the Southern Hemisphere during the Permian-Triassic transition.The results show that the foraminifer of the Selong Section experienced a two-pulsed extinction(total species extinction rate of 71%),consistent with the time in South China but with a lower magnitude of extinction.Meanwhile,the data show that foraminiferal test volume was significantly miniaturized following the first pulse of extinction event:the mean size of post-extinction foraminifer was only 15%of that in the pre-extinction,mainly reflected by the disappearance of large forms as well as occurrences of smaller survivors and originators.Combined with the South China record,size data from southern Tibet indicate that the miniaturization of foraminifera is synchronous in the Paleotethys and Neotethys but smaller in magnitude in the Neotethys.We propose that ocean anoxia and acidification may be the environmental pressures leading to local and global foraminiferal miniaturizations,along with global warming,which might play a dominant role.展开更多
Extinction selectivity determines the direction of macroevolution,especially during mass extinction;however,its driving mechanisms remain poorly understood.By investigating the physiological selectivity of marine anim...Extinction selectivity determines the direction of macroevolution,especially during mass extinction;however,its driving mechanisms remain poorly understood.By investigating the physiological selectivity of marine animals during the Permian-Triassic mass extinction,we found that marine clades with lower O2-carrying capacity hemerythrin proteins and those relying on O2 diffusion experienced significantly greater extinction intensity and body-size reduction than those with higher O2-carrying capacity hemoglobin or hemocyanin proteins.Our findings suggest that animals with high O2-carrying capacity obtained the necessary O2 even under hypoxia and compensated for the increased energy requirements caused by ocean acidification,which enabled their survival during the Permian-Triassic mass extinction.Thus,high O2-carrying capacity may have been crucial for the transition from the Paleozoic to the Modern Evolutionary Fauna.展开更多
Climatic and environmental conditions play a pivotal role in the evolution of the biosphere,serving as the primary natural factors influencing biological evolution and the development of human civilization.The study o...Climatic and environmental conditions play a pivotal role in the evolution of the biosphere,serving as the primary natural factors influencing biological evolution and the development of human civilization.The study of the evolution of Earth's habitability primarily revolves around the reconstruction of climatic and oceanic conditions in geohistorical periods,shedding light on their dynamic changes.This paper collates classic geological indicators and geochemical proxies associated with paleoclimatic and oceanic environmental conditions.The latest“big data”analyses and simulations made possible by the availability of previously unimagined massive datasets reveal several key findings:During the early Paleozoic,atmospheric oxygen levels were low,and widespread oceanic anoxia was prevalent;the Devonian era witnessed a greenhouse climate,followed by the Carboniferous ice age characterized by higher oceanic oxidation levels and alkalinity.The latest Paleozoic deglaciation occurred under high pCO_(2) conditions,extending into much of the Mesozoic and early Cenozoic,marked by multiple hyperthermal and anoxia expansion events,until the resurgence of global glaciation in the middle-late stages of the Cenozoic,ultimately bringing environmental and climatic conditions closer to modern levels.By correlating the aforementioned long-term trends with major geological events,we can delineate the co-evolution of paleoclimate and oceanic environments in tandem with the development of Tethys tectonics as follows.(1)During the Proto-Tethys stage,global paleo-elevations were relatively low,and atmospheric oxygen levels were also relatively modest.Despite the occurrence of significant tectonic movements that led to noticeable transgressive-regressive cycles,their effects on climate and oceanic environments were somewhat limited due to the relatively weak interactions.(2)The emergence of the Paleo-Tethys was a significant event that coincided with the formation of the supercontinent Pangaea.Intensive orogenic movements during this 展开更多
Various environmental changes were associated with the Permian-Triassic mass extinction at 252.2 Ma. Diverse unusual sediments and depositional phenomena have been uncovered as responses to environmental and biotic ch...Various environmental changes were associated with the Permian-Triassic mass extinction at 252.2 Ma. Diverse unusual sediments and depositional phenomena have been uncovered as responses to environmental and biotic changes. Lithological and detailed conodont biostratigraphic correlations within six Permian-Triassic boundary sections in South China indicate rapid fluctuations in carbonate deposition. Four distinct depositional phases can be recognized： （1） normal carbonate deposition on the platform and slope during the latest Permian; （2） reduced carbonate deposition at the on- set of the main extinction horizon; （3） expanded areas of carbonate deposition during the Hindeodus changxingsensis Zone to the H. parvus Zone; and （4） persistent mud-enriched carbonate deposition in the aftermath of the Permian-Triassic transition. Although availability of skeletal carbonate was significantly reduced during the mass extinction, the increase in carbonate deposition did not behave the same way. The rapid carbonate depositional changes, presented in this study, suggest that diverse environmental changes played key roles in the carbonate deposition of the Permian-Triassic mass extinction and onset of its aftermath. An overview of hypotheses to explain these changes implies enhanced terrestrial input, abnormal ocean circulation and various geobiological processes contributed to carbonate saturation fluctuations, as the sedimentary response to large volcanic eruptions.展开更多
Ocean anoxia has been widely implicated in the Permian-Triassic extinction. However, the duration and distribution of the ocean anoxia remains controversial. In this study, the detailed redox changes across the Permia...Ocean anoxia has been widely implicated in the Permian-Triassic extinction. However, the duration and distribution of the ocean anoxia remains controversial. In this study, the detailed redox changes across the Permian-Triassic boundary （PTB） in the shallow platform interior at Great Bank of Guizhou （GBG） has been reconstructed based on the high-resolution microfossil composition and multiple paleo-redox proxies. The shallow platform is characterized by low sulfur （total sulfur （TS） and pyrite sulfur （Spy）） concentrations, low Spy/TOC ratios, and low DOP values before the mass extinction, representing oxic conditions well. Following the mass extinction, the shift of multiple geochemical proxies, including high Spy/TOC ratios and DOP values, indicates dysoxic-anoxic conditions in shallow ocean. Furthermore, we reconstruct the transition of the redox conditions of Nanpanjiang Basin： the intense volcanic eruptions, which release huge COz and SO2 before the mass extinction, provoke the temperature rising and the collapse of terrestrial ecosystem. As a result, the increased weathering influx causes the carbon iso- topic negative excursion and the expansion of the ocean oxygen minimum zone （OMZ）. When the OMZ expanded into the photic zone, the episodic H2S release events enhance the pyrite burial at Dajiang section. Thus, intense volcanic eruptions, temperature increase, and oceanic hypoxia together lead to the PTB extinction. Recent studies show high temperature might be the key mechanism of the PTB extinction. In addition, this study confirms that the microbialites were formed in the dysoxic- anoxic shallow water.展开更多
基金supported by the National Natural Science Foundation of China (Grant Nos. 41530104 & 41661134047)
文摘The Triassic rocks are widespread in China, and both marine and terrestrial strata are well developed. The Triassic stratigraphic architecture of China is very complex in both spatial variation of the so-called "South Marine and North Continental", i.e. the southern areas of China occupied mostly by marine facies while the northern China by terrestrial facies during the Triassic Period, and temporal transition of the "Lower Marine and Upper Continental", i.e. the lower part of the Triassic System composed mainly of marine facies and the upper part of terrestrial strata especially in South China. Although the Global Stratotype Section and Point(GSSP) of the Permian-Triassic boundary is located in South China, the Triassic of China except for some marine Lower-Middle Triassic depositions shows significantly local characteristics and is hardly correlated with the global chronostratigraphic chart. Consequently, the Triassic of China contains not only the international research hotspots but also difficult points in stratigraphic study. This paper aims to present a brief review of the Triassic in China, including chronostratigraphy, biostratigraphy, magnetostratigraphy and chemostratigraphy, and summarize an integrated Triassic stratigraphic framework of China. Accordingly, a stratigraphic correlation is proposed for the lithostratigraphic sequences among the three tectono-paleogeographic stratigraphic regions. The comprehensive study indicates that ammonoids are the classic index fossils in Triassic biostratigraphy but conodonts are more advantageous in the study and definition of the Triassic chronostratigraphic boundaries. China still has the potential to optimize the GSSPs of the Induan-Olenekian boundary and Olenekian-Anisian boundary. The correlation of the Permian-Triassic boundary between marine and terrestrial facies might be achieved with the help of the Permian-Triassic "transitional bed" and its related biotic and environmental events in association with the biostratigraphic study of conchostracan, vertebra
基金supported by the National Natural Science Foundation of China (Nos. 41821001, 41622207, 41530104, 41661134047)the State Key R&D Project of China (No. 2016YFA0601100)+3 种基金the Strategic Priority Research Program of Chinese Academy of Sciences (No. XDB26000000)a Marie Curie Fellowship (No. H2020MSCA-IF-2015-701652)the Natural Environment Research Council’s Eco-PT Project (No. NE/P01377224/1)a part of the Biosphere Evolution, Transitions and Resilience Program (BETR)
文摘Ocean temperature and dissolved oxygen concentrations are critical factors that control ocean productivity, carbon and nutrient cycles, and marine habitat. However, the evolution of these two factors in the geologic past are still unclear. Here, we use a new oxygen isotope database to establish the sea surface temperature(SST) curve in the past 500 million years. The database is composed of 22 796 oxygen isotope values of phosphatic and calcareous fossils. The result shows two prolonged cooling events happened in the Late Paleozoic and Late Cenozoic, coinciding with two major ice ages indicated by continental glaciation data, and seven global warming events that happened in the Late Cambrian, Silurian–Devonian transition, Late Devonian, Early Triassic, Toarcian, Late Cretaceous, and Paleocene–Eocene transition. The SSTs during these warming periods are about 5–30 °C higher than the present-day level. Oxygen contents of shallow seawater are calculated from temperature, salinity, and atmospheric oxygen. The results show that major dissolved oxygen valleys of surface seawater coincide with global warming events and ocean anoxic events. We propose that the combined effect of temperature and dissolved oxygen account for the long-term evolution of global oceanic redox state during the Phanerozoic.
基金supported by the State Key R&D Project of China(No.2023YFF0804000)the National Natural Science Foundation of China(Nos.92155201,92255303)a BQR of the Universit'e de Bourgogne。
文摘The miniaturization of organisms during the Permian-Triassic mass extinction,as an ecological strategy in response to environmental devastation,has been widely recognized in diverse marine invertebrates.Previous studies on the extinction process and miniaturization of foraminifers in the Permian-Triassic interval have relied on the fossil record of the low-latitude Paleotethys or a global database,although data and materials from the high-latitude Neotethys region are still rare.To reveal the evolutionary patterns and spatial variability of foraminifers at different latitudes and paleogeographic contexts,here we investigated the fossil distribution and size variation of foraminifers in the Selong Section of southern Tibet,located in the mid-latitude Neotethys of the Southern Hemisphere during the Permian-Triassic transition.The results show that the foraminifer of the Selong Section experienced a two-pulsed extinction(total species extinction rate of 71%),consistent with the time in South China but with a lower magnitude of extinction.Meanwhile,the data show that foraminiferal test volume was significantly miniaturized following the first pulse of extinction event:the mean size of post-extinction foraminifer was only 15%of that in the pre-extinction,mainly reflected by the disappearance of large forms as well as occurrences of smaller survivors and originators.Combined with the South China record,size data from southern Tibet indicate that the miniaturization of foraminifera is synchronous in the Paleotethys and Neotethys but smaller in magnitude in the Neotethys.We propose that ocean anoxia and acidification may be the environmental pressures leading to local and global foraminiferal miniaturizations,along with global warming,which might play a dominant role.
基金State Key R&D Project of China(2023YFF0804000)National Natural Science Foundation of China(42325202,92155201,and 92255303),the 111 Project(B08030)+1 种基金Natural Science Foundation of Hubei(2023 AFA006)Fundamental Research Funds for the Central Universities,China University of Geosciences(Wuhan).
文摘Extinction selectivity determines the direction of macroevolution,especially during mass extinction;however,its driving mechanisms remain poorly understood.By investigating the physiological selectivity of marine animals during the Permian-Triassic mass extinction,we found that marine clades with lower O2-carrying capacity hemerythrin proteins and those relying on O2 diffusion experienced significantly greater extinction intensity and body-size reduction than those with higher O2-carrying capacity hemoglobin or hemocyanin proteins.Our findings suggest that animals with high O2-carrying capacity obtained the necessary O2 even under hypoxia and compensated for the increased energy requirements caused by ocean acidification,which enabled their survival during the Permian-Triassic mass extinction.Thus,high O2-carrying capacity may have been crucial for the transition from the Paleozoic to the Modern Evolutionary Fauna.
基金supported by the National Natural Science Foundation of China (Grant Nos.92155201,42272361)China Geological Survey (The Establishment of Chinese Stratigraphic Standards Project)。
文摘Climatic and environmental conditions play a pivotal role in the evolution of the biosphere,serving as the primary natural factors influencing biological evolution and the development of human civilization.The study of the evolution of Earth's habitability primarily revolves around the reconstruction of climatic and oceanic conditions in geohistorical periods,shedding light on their dynamic changes.This paper collates classic geological indicators and geochemical proxies associated with paleoclimatic and oceanic environmental conditions.The latest“big data”analyses and simulations made possible by the availability of previously unimagined massive datasets reveal several key findings:During the early Paleozoic,atmospheric oxygen levels were low,and widespread oceanic anoxia was prevalent;the Devonian era witnessed a greenhouse climate,followed by the Carboniferous ice age characterized by higher oceanic oxidation levels and alkalinity.The latest Paleozoic deglaciation occurred under high pCO_(2) conditions,extending into much of the Mesozoic and early Cenozoic,marked by multiple hyperthermal and anoxia expansion events,until the resurgence of global glaciation in the middle-late stages of the Cenozoic,ultimately bringing environmental and climatic conditions closer to modern levels.By correlating the aforementioned long-term trends with major geological events,we can delineate the co-evolution of paleoclimate and oceanic environments in tandem with the development of Tethys tectonics as follows.(1)During the Proto-Tethys stage,global paleo-elevations were relatively low,and atmospheric oxygen levels were also relatively modest.Despite the occurrence of significant tectonic movements that led to noticeable transgressive-regressive cycles,their effects on climate and oceanic environments were somewhat limited due to the relatively weak interactions.(2)The emergence of the Paleo-Tethys was a significant event that coincided with the formation of the supercontinent Pangaea.Intensive orogenic movements during this
基金supported by the "973 Program" (No. 2011CB808800)the National Natural Science Foundation of China (Nos. 41172312, 41272372, 41302010, 41402302)+1 种基金the State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences (No. GKZ14Y663)the "111 Project" (No. B08030)
文摘Various environmental changes were associated with the Permian-Triassic mass extinction at 252.2 Ma. Diverse unusual sediments and depositional phenomena have been uncovered as responses to environmental and biotic changes. Lithological and detailed conodont biostratigraphic correlations within six Permian-Triassic boundary sections in South China indicate rapid fluctuations in carbonate deposition. Four distinct depositional phases can be recognized： （1） normal carbonate deposition on the platform and slope during the latest Permian; （2） reduced carbonate deposition at the on- set of the main extinction horizon; （3） expanded areas of carbonate deposition during the Hindeodus changxingsensis Zone to the H. parvus Zone; and （4） persistent mud-enriched carbonate deposition in the aftermath of the Permian-Triassic transition. Although availability of skeletal carbonate was significantly reduced during the mass extinction, the increase in carbonate deposition did not behave the same way. The rapid carbonate depositional changes, presented in this study, suggest that diverse environmental changes played key roles in the carbonate deposition of the Permian-Triassic mass extinction and onset of its aftermath. An overview of hypotheses to explain these changes implies enhanced terrestrial input, abnormal ocean circulation and various geobiological processes contributed to carbonate saturation fluctuations, as the sedimentary response to large volcanic eruptions.
基金supported by National Basic Research Program of China (Grant No. 2011CB808800)National Natural Science Foundation of China (Grant Nos. 41172312, 41272372, 41172036, 41240016, 41302271)+1 种基金Fundamental Research Funds for the Central Universities, China University of Geosciences (Wuhan)Fund of State Key Laboratory of Biogeology and Environmental Geology (Grant No. BGEG1016)
文摘Ocean anoxia has been widely implicated in the Permian-Triassic extinction. However, the duration and distribution of the ocean anoxia remains controversial. In this study, the detailed redox changes across the Permian-Triassic boundary （PTB） in the shallow platform interior at Great Bank of Guizhou （GBG） has been reconstructed based on the high-resolution microfossil composition and multiple paleo-redox proxies. The shallow platform is characterized by low sulfur （total sulfur （TS） and pyrite sulfur （Spy）） concentrations, low Spy/TOC ratios, and low DOP values before the mass extinction, representing oxic conditions well. Following the mass extinction, the shift of multiple geochemical proxies, including high Spy/TOC ratios and DOP values, indicates dysoxic-anoxic conditions in shallow ocean. Furthermore, we reconstruct the transition of the redox conditions of Nanpanjiang Basin： the intense volcanic eruptions, which release huge COz and SO2 before the mass extinction, provoke the temperature rising and the collapse of terrestrial ecosystem. As a result, the increased weathering influx causes the carbon iso- topic negative excursion and the expansion of the ocean oxygen minimum zone （OMZ）. When the OMZ expanded into the photic zone, the episodic H2S release events enhance the pyrite burial at Dajiang section. Thus, intense volcanic eruptions, temperature increase, and oceanic hypoxia together lead to the PTB extinction. Recent studies show high temperature might be the key mechanism of the PTB extinction. In addition, this study confirms that the microbialites were formed in the dysoxic- anoxic shallow water.
