基金supported by the National Natural Science Foundation of China(No.41790455,41772008)the National Geographic Society,USA(No.EC0768-15)+1 种基金the Fundamental Research Funds for the Central Universities(No.2652017215)the State Key Laboratory of Palaeobiology and Stratigraphy(Nanjing Institute of Geology and Palaeontology,Chinese Academy of Sciences)(No.173127)