基金HaK was supported by the National Natural Science Foundation of China(92043301,82030103,and 91843302)and China Medical Board Collaborating Program(16-250)AG and FS were supported by the Medical Research Council,UK(MK/M022625/1)+10 种基金the Natural Environment Research Council,UK(NE/R009384/1)and the European Union's Horizon 2020 Project Exhaustion(820655)VH was supported by the Spanish Ministry of Science and Innovation(PCIN-2017-046)and the German Federal Ministry of Education and Research(01 LSI201A2)YH and MH were supported by the Environment Research and Technology Development Fund(JPMEERF15S11412)the Environmental Restoration and Conservation Agency,Japan.JK and AU were supported by the Czech Science Foundation(18-22125S)ST was supported by the Shanghai Municipal Science and Technology Commission(18411951600)Y-LLG was supported by a Career Development Fellowship of the Australian National Health and Medical Research Council(APP1163693)SL was supported by an Early Career Fellowship of the Australian National Health and Medical Research Council(APP1109193)JJKJJ and NR were supported by the Academy of Finland(310372)The funders had no role in considering the study design or in the collection,analysis,interpretation of data,writing of the report,or decision to submit the article for publication.